01 - Preheat the oven to 375°F. Line a 12-cup muffin tin with paper liners or grease thoroughly.
02 - In a large bowl, whisk together flour, sugar, salt, baking powder, and baking soda until well blended.
03 - In a separate bowl, whisk together sourdough starter, eggs, melted butter, milk, and vanilla extract until smooth and homogeneous.
04 - Pour the wet mixture into the dry ingredients. Gently fold together until just combined, being careful not to overmix.
05 - Gently fold in the blueberries and lemon zest if using, distributing evenly throughout the batter.
06 - Divide the batter evenly among the prepared muffin cups, filling each about three-quarters full.
07 - Bake for 22 to 25 minutes, or until a toothpick inserted into the center comes out clean and the tops are golden brown.
08 - Let the muffins cool in the pan for 5 minutes, then transfer to a wire rack to cool completely.